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The Story of Evolution
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there was no other conceivable source that would supply the sun's tremendous outpour of energy for tens of millions of years except the compression of its substance. It is true that the discovery of radio-activity has disclosed a new source of energy within the atoms themselves, and there are scientific men, like Professor Arrhenius, who attach great importance to this source. But, although it may prolong the limited term of life which physicists formerly allotted to the sun and other stars, it is still felt that the condensation of a nebula offers the best explanation of the origin of a sun, and we have ample evidence for the connection. We must, therefore, see what the nebula is, and how it develops.

"Nebula" is merely the Latin word for cloud. Whatever the nature of these diffused stretches of matter may be, then, the name applies fitly to them, and any theory of the development of a star from them is still a "nebular hypothesis." But the three theories which divide astronomers to-day differ as to the nature of the nebula. The older theory, pointing to the gaseous nebulae as the first stage, holds that the nebula is a cloud of extremely attenuated gas. The meteoritic hypothesis (Sir N. Lockyer, Sir G. Darwin, etc.), observing that space seems to swarm with meteors and that the greater part of the nebulae are not gaseous, believes that the starting-point is a colossal swarm of meteors, surrounded by the gases evolved and lit up by their collisions. The planetesimal hypothesis, advanced in recent years by Professor Moulton and Professor Chamberlin, contends that the nebula is a vast cloud of liquid or solid (but not gaseous) particles. This theory is based mainly on the dynamical difficulties of the other two, which we will notice presently.

The truth often lies between conflicting theories, or they may apply to different cases. It is not improbable that this will be our experience in regard to the nature of the initial nebula. The gaseous nebulae, and the formation of such nebulae from disrupted stars, are facts that cannot be ignored. The nebulae with a continuous spectrum, and therefore--in part, at least--in a liquid or solid condition, may very well be regarded as a more advanced stage of condensation of the same; their spiral shape and conspicuous nuclei are consistent with this. Moreover, a condensing swarm of meteors would, owing to the heat evolved, tend to pass into a gaseous condition. On the tether hand, a huge expanse of gas stretched over billions of miles of space would be a net for the wandering particles, meteors, and comets that roam through space. If it be true, as is calculated, that our 24,000 miles of atmosphere capture a hundred million meteors a day, what would the millions or billions of times larger net of a nebula catch, even if the gas is so much thinner? In other words, it is not wise to draw too fine a line between a gaseous nebula and one consisting of solid particles with gas.

The more important question is: How do astronomers conceive the condensation of this mixed mass of cosmic dust? It is easy to reply that gravitation, or the pressure of the surrounding ether, slowly drives the particles centre-ward, and compresses the dust into globes, as the boy squeezes the flocculent snow into balls; and it is not difficult for the mathematician to show that this condensation would account for the shape and temperature of the stars. But we must go a little beyond this superficial statement, and see, to some extent, how the deeper students work out the process.*

* See, especially, Dr. P. Lowell, "The Evolution of Worlds" (1909). Professor S. Arrhenius, "Worlds in the Making" (1908), Sir N. Lockyer, "The Meteorite Hypothesis" (1890), Sir R. Ball, "The Earth's Beginning" (1909), Professor Moulton, "The Astrophysical Journal (October, 1905), and Chamberlin and Salisbury, "Geology," Vol. II. (1903).



Taking a broad view of the whole field, one may say that the two chief difficulties are as follows: First, how to get the whole chaotic mass whirling round in one common direction; secondly, how to account for the fact that in our solar system the outermost planets and satellites do not rotate in the same direction as the rest. There is a widespread idea that these difficulties have proved fatal to the old nebular hypothesis, and there are distinguished astronomers who think so. But Sir R. Ball (see note), Professor Lowell (see note), Professor Pickering (Annals of Harvard College Observatory, 53, III), and other high authorities deny this, and work out the newly discovered movements on the lines of the old theory. They hold that all the bodies in the solar system once turned in the same direction as Uranus and Neptune, and the tidal influence of the sun has changed the rotation of most of them. The planets farthest from the sun would naturally not be so much affected by it. The same principle would explain the retrograde movement of the outer satellites of Saturn and Jupiter. Sir R. Ball further works out the principles on which the particles of the condensing nebula would tend to form a disk rotating on its central axis. The ring-theory of Laplace is practically abandoned. The spiral nebula is evidently the standard type, and the condensing nebula must conform to it. In this we are greatly helped by the current theory of the origin of spiral nebulae.

We saw previously that new stars sometimes appear in the sky, and the recent closer scrutiny of the heavens shows this occurrence to be fairly frequent. It is still held by a few astronomers that such a cataclysm means that two stars collided. Even a partial or "grazing " collision between two masses, each weighing billions of tons, travelling (on the average) forty or fifty miles a second--a movement that would increase enormously as they approach each other--would certainly liquefy or vaporise their substance; but the astronomer, accustomed to see cosmic bodies escape each other by increasing their speed, is generally disinclined to believe in collisions. Some have made the new star plunge into the heart of a dense and dark nebula; some have imagined a shock of two gigantic swarms of meteors; some have regarded the outflame as the effect of a prodigious explosion. In one or other new star each or any of these things may have occurred, but the most plausible and accepted theory for the new star of 1901 and some others is that two stars had approached each other too closely in their wandering. Suppose that, in millions of years to come, when our sun is extinct and a firm crust surrounds the great molten ball, some other sun approaches within a few million miles of it. The two would rush past each other at a terrific speed, but the gravitational effect of the approaching star would tear open the solid shell of the sun, and, in a mighty flame, its molten and gaseous entrails would be flung out into space. It has long been one of the arguments against a molten interior of the earth that the sun's gravitational influence would raise it in gigantic tides and rend the solid shell of rock. It is even suspected now that our small earth is not without a tidal influence on the sun. The comparatively near approach of two suns would lead to a terrific cataclysm.

If we accept this theory, the origin of the spiral nebula becomes intelligible. As the sun from which it is formed is already rotating on its axis, we get a rotation of the nebula from the first. The mass poured out from the body of the sun would, even if it were only a small fraction of its mass, suffice to make a planetary system; all our sun's planets and their satellites taken together amount to only 1/100th of the mass of the solar system. We may assume, further, that the outpoured matter would be a mixed cloud of gases and solid and liquid particles; and that it would stream out, possibly in successive waves, from more than one part of the disrupted sun, tending to form great spiral trails round the parent mass. Some astronomers even suggest that, as there are tidal waves raised by the moon at opposite points of the earth, similar tidal outbursts would occur at opposite points on the disk of the disrupted star, and thus give rise to the characteristic arms starting from opposite sides of the spiral nebula. This is not at all clear, as the two tidal waves of the earth are due to the fact that it has a liquid ocean rolling on, not under, a solid bed.

In any case, we have here a good suggestion of the origin of the spiral nebula and of its further development. As soon as the outbursts are over, and the scattered particles have reached the farthest limit to which they are hurled, the concentrating action of gravitation will slowly assert itself. If we conceive this gravitational influence as the pressure of the surrounding ether we get a wider understanding of the process. Much of the dispersed matter may have been shot far enough into space to escape the gravitational pull of the parent mass, and will be added to the sum of scattered cosmic dust, meteors, and close shoals of meteors (comets) wandering in space. Much of the rest will fall back upon the central body But in the great spiral arms themselves the distribution of the matter will be irregular, and the denser areas will slowly gather in the surrounding material. In the end we would thus get secondary spheres circling round a large primary.

This is the way in which astronomers now generally conceive the destruction and re-formation of worlds. On one point the new planetesimal theory differs from the other theories. It supposes that, since the particles of the whirling nebula are all travelling in the same general direction, they overtake each other with less violent impact than the other theories suppose, and therefore the condensation of the material into planets would not give rise to the terrific heat which is generally assumed. We will consider this in the next chapter, when we deal with the formation of the planets. As far as the central body, the sun, is concerned, there can be no hesitation. The 500,000,000 incandescent suns in the heavens are eloquent proof of the appalling heat that is engendered by the collisions of the concentrating particles.

In general outline we now follow the story of a star with some confidence. An internal explosion, a fatal rush into some dense nebula or swarm of meteors, a collision with another star, or an approach within a few million miles of another star, scatters, in part or whole, the solid or liquid globe in a cloud of cosmic dust. When the violent outrush is over, the dust is gathered together once more into a star. At first cold and attenuated, its temperature rises as the particles come together, and we have, after a time, an incandescent nucleus shining through a thin veil of gas--a nebulous star. The temperature rises still further, and we have the blue-hot star, in which the elements seem to be dissociated, and slowly re-forming as the temperature falls. After, perhaps, hundreds of millions of years it reaches the "yellow" stage, and, if it has planets with the conditions of life, there may be a temporary opportunity for living things to enjoy its tempered energy. But the cooler vapours are gathering round it, and at length its luminous body is wholly imprisoned. It continues its terrific course through space, until some day, perhaps, it again encounters the mighty cataclysm which will make it begin afresh the long and stormy chapters of its living history.

Such is the suggestion of the modern astronomer, and, although we seem to find every phase of the theory embodied in the varied contents of the heavens, we must not forget that it is only a suggestion. The spectroscope and telescopic photography, which are far more important than the visual telescope, are comparatively recent, and the field to be explored is enormous. The mist is lifting from the cosmic landscape, but there is still enough to blur our vision. Very puzzling questions remain unanswered. What is the origin of the great gaseous nebulae? What is the origin of the triple or quadruple star? What is the meaning of stars whose light ebbs and flows in periods of from a few to several hundred days? We may even point to the fact that some, at least, of the spiral nebulae are far too vast to be the outcome of the impact or approach of two stars.

We may be content to think that we have found out some truths, by no means the whole truth, about the evolution of worlds. Throughout this immeasurable ocean of ether the particles of matter are driven together and form bodies. These bodies swarm throughout space, like fish in the sea; travelling singly (the "shooting star"), or in great close shoals (the nucleus of a comet), or lying scattered in vast clouds. But the inexorable pressure urges them still, until billions of tons of material are gathered together. Then, either from the sheer heat of the compression, or from the formation of large and unstable atomic systems (radium, etc.), or both, the great mass becomes a cauldron of fire, mantled in its own vapours, and the story of a star is run. It dies out in one part of space to begin afresh in another. We see nothing in the nature of a beginning or an end for the totality of worlds, the universe. The life of all living things on the earth, from the formation of the primitive microbes to the last struggles of the superman, is a small episode of that stupendous drama, a fraction of a single scene. But our ampler knowledge of it, and our personal interest in it, magnify that episode, and we turn from the cosmic picture to study the formation of the earth and the rise of its living population.





CHAPTER IV. THE PREPARATION OF THE EARTH

The story of the evolution of our solar system is, it will now be seen, a local instance of the great cosmic process we have studied in the last chapter. We may take one of the small spiral nebulae that abound in the heavens as an illustration of the first stage. If a still earlier stage is demanded, we may suppose that some previous sun collided with, or approached too closely, another mighty body, and belched out a large part of its contents in mighty volcanic outpours. Mathematical reasoning can show that this erupted material would gather into a spiral nebula; but, as mathematical calculations cannot be given here, and are less safe than astronomical facts, we will be content to see the early shape of our solar system in a relatively small spiral nebula, its outermost arm stretching far beyond the present orbit of Neptune, and its great nucleus being our present sun in more diffused form.

We need not now attempt to follow the shrinking of the central part of the nebula until it becomes a rounded fiery sun. That has been done in tracing the evolution of a star. Here we have to learn how the planets were formed from the spiral arms of the nebula. The principle of their formation is already clear. The same force of gravitation, or the same pressure of the surrounding ether, which compresses the central mass into a fiery globe, will act upon the loose material of the arms and compress it into smaller globes. But there is an interesting and acute difference of opinion amongst modern experts as to whether these smaller globes, the early planets, would become white-hot bodies.

The general opinion, especially among astronomers, is that the compression of the nebulous material of the arms into globes would generate enormous heat, as in the case of the sun. On that view the various planets would begin their careers as small suns, and would pass through those stages of cooling and shrinking which we have traced in the story of the stars. A glance at the photograph of one of the spiral nebulae strongly confirms this. Great luminous knots, or nuclei, are seen at intervals in the arms. Smaller suns seem to be forming in them, each gathering into its body the neighbouring material of the arm, and rising in temperature as the mass is compressed into a globe. The spectroscope shows that these knots are condensing masses of white-hot liquid or solid matter. It therefore seems plain that each planet will first become a liquid globe of fire, coursing round the central sun, and will gradually, as its heat is dissipated and the supply begins to fail, form a solid crust.

This familiar view is challenged by the new "planetesimal hypothesis," which has been adopted by many distinguished geologists (Chamberlin, Gregory, Coleman, etc.). In their view the particles in the arms of the nebula are all moving in the same direction round the sun. They therefore quietly overtake the nucleus to which they are attracted, instead of violently colliding with each other, and much less heat is generated at the surface. In that case the planets would not pass through a white-hot, or even red-hot, stage at all. They are formed by a slow ingathering of the scattered particles, which are called "planetesimals" round the larger or denser masses of stuff which were discharged by the exploding sun. Possibly these masses were prevented from falling back into the sun by the attraction of the colliding body, or the body which caused the eruption. They would revolve round the parent body, and the shoals of smaller particles would gather about them by gravitation. If there were any large region in the arm of the nebula which had no single massive nucleus, the cosmic dust would gather about a number of smaller centres. Thus might be explained the hundreds of planetoids, or minor planets, which we find between Mars and Jupiter. If these smaller bodies came within the sphere of influence of one of the larger planets, yet were travelling quickly enough to resist its attraction, they would be compelled to revolve round it, and we could thus explain the ten satellites of Saturn and the eight of Jupiter. Our moon, we shall see, had a different origin.

We shall find this new hypothesis crossing the familiar lines at many points in the next few chapters. We will consider those further consequences as they arise, but may say at once that, while the new theory has greatly helped us in tracing the formation of the planetary system, astronomers are strongly opposed to its claim that the planets did not pass through an incandescent stage. The actual features of our spiral nebulae seem clearly to exhibit that stage. The shape of the planets--globular bodies, flattened at the poles--strongly suggests that they were once liquid. The condition in which we find Saturn and Jupiter very forcibly confirms this suggestion; the latest study of those planets supports the current opinion that they are still red-hot, and even seems to detect the glow of their surfaces in their mantles of cloud. These points will be considered more fully presently. For the moment it is enough to note that, as far as the early stages of planetary development are concerned, the generally accepted theory rests on a mass of positive evidence, while the new hypothesis is purely theoretical. We therefore follow the prevailing view with some confidence.

Those of the spiral nebulae which face the earth squarely afford an excellent suggestion of the way in which planets are probably formed. In some of these nebulae the arms consist of almost continuous streams of faintly luminous matter; in others the matter is gathering about distinct centres; in others again the nebulous matter is, for the most part, collected in large glowing spheres. They seem to be successive stages, and to reveal to us the origin of our planets. The position of each planet in our solar system would be determined by the chance position of the denser stuff shot out by the erupting sun. I have seen Vesuvius hurl up into the sky, amongst its blasts of gas and steam, white-hot masses of rock weighing fifty tons. In the far fiercer outburst of the erupting sun there would be at least thinner and denser masses, and they must have been hurled so far into space that their speed in travelling round the central body, perhaps seconded by the attraction of the second star, overcame the gravitational pull back to the centre. Recollect the force which, in the new star in Perseus, drove masses of hydrogen for millions of miles at a speed of a thousand miles a second.

These denser nuclei or masses would, when the eruption was over, begin to attract to themselves all the lighter nebulous material within their sphere of gravitational influence. Naturally, there would at first be a vast confusion of small and large centres of condensation in the arms of the nebula, moving in various directions, but a kind of natural selection--and, in this case, survival of the biggest--would ensue. The conflicting movements would be adjusted by collisions and gravitation, the smaller bodies would be absorbed in the larger or enslaved as their satellites, and the last state would be a family of smaller suns circling at vast distances round the parent body. The planets, moreover, would be caused to rotate on their axes, besides revolving round the sun, as the particles at their inner edge (nearer the sun) would move at a different speed from those at the outer edge. In the course of time the smaller bodies, having less heat to lose and less (or no) atmosphere to check the loss, would cool down, and become dark solid spheres, lit only by the central fire.

While the first stage of this theory of development is seen in the spiral nebula, the later stages seem to be well exemplified in the actual condition of our planets. Following, chiefly, the latest research of Professor Lowell and his colleagues, which marks a considerable advance on our previous knowledge, we shall find it useful to glance at the sister-planets before we approach the particular story of our earth.

Mercury, the innermost and smallest of the planets, measuring only some 3400 miles in diameter, is, not unexpectedly, an airless wilderness. Small bodies are unable to retain the gases at their surface, on account of their feebler gravitation. We find, moreover, that Mercury always presents the same face to the sun, as it turns on its axis in the same period (eighty-eight days) in which it makes a revolution round the sun. While, therefore, one half of the globe is buried in eternal darkness, the other half is eternally exposed to the direct and blistering rays of the sun, which is only 86,000,000 miles away. To Professor Lowell it presents the appearance of a bleached and sun-cracked desert, or "the bones of a dead world." Its temperature must be at least 300 degrees C. above that of the earth. Its features are what we should expect on the nebular hypothesis. The slowness of its rotation is accounted for by the heavy tidal influence of the sun. In the same way our moon has been influenced by the earth, and our earth by the sun, in their movement of rotation.

Venus, as might be expected in the case of so large a globe (nearly as large as the earth), has an atmosphere, but it seems, like Mercury, always to present the same face to the sun. Its comparative nearness to the sun (67,000,000 miles) probably explains this advanced effect of tidal action. The consequences that the observers deduce from the fact are interesting. The sun-baked half of Venus seems to be devoid of water or vapour, and it is thought that all its water is gathered into a rigid ice-field on the dark side of the globe, from which fierce hurricanes must blow incessantly. It is a Sahara, or a desert far hotter than the Sahara, on one side; an arctic region on the other. It does not seem to be a world fitted for the support of any kind of life that we can imagine.

When we turn to the consideration of Mars, we enter a world of unending controversy. With little more than half the diameter of the earth, Mars ought to be in a far more advanced stage of either life or decay, but its condition has not yet been established. Some hold that it has a considerable atmosphere; others that it is too small a globe to have retained a layer of gas. Professor Poynting believes that its temperature is below the freezing-point of water all over the globe; many others, if not the majority of observers, hold that the white cap we see at its poles is a mass of ice and snow, or at least a thick coat of hoar-frost, and that it melts at the edges as the springtime of Mars comes round. In regard to its famous canals we are no nearer agreement. Some maintain that the markings are not really an objective feature; some hold that they are due to volcanic activity, and that similar markings are found on the moon; some believe that they are due to clouds; while Professor Lowell and others stoutly adhere to the familiar view that they are artificial canals, or the strips of vegetation along such canals. The question of the actual habitation of Mars is still open. We can say only that there is strong evidence of its possession of the conditions of life in some degree, and that living things, even on the earth, display a remarkable power of adaptation to widely differing conditions.

Passing over the 700 planetoids, which circulate between Mars and Jupiter, and for which we may account either by the absence of one large nucleus in that part of the nebulous stream or by the disturbing influence of Jupiter, we come to the largest planet of the system. Here we find a surprising confirmation of the theory of planetary development which we are following. Three hundred times heavier than the earth (or more than a trillion tons in weight), yet a thousand times less in volume than the sun, Jupiter ought, if our theory is correct, to be still red-hot. All the evidence conspires to suggest that it is. It has long been recognised that the shining disk of the planet is not a solid, but a cloud, surface. This impenetrable mass of cloud or vapour is drawn out in streams or belts from side to side, as the giant globe turns on its axis once in every ten hours. We cannot say if, or to what extent, these clouds consist of water-vapour. We can conclude only that this mantle of Jupiter is "a seething cauldron of vapours" (Lowell), and that, if the body beneath is solid, it must be very hot. A large red area, at one time 30,000 miles long, has more or less persisted on the surface for several decades, and it is generally interpreted, either as a red-hot surface, or as a vast volcanic vent, reflecting its glow upon the clouds. Indeed, the keen American observers, with their powerful telescopes, have detected a cherry-red glow on the edges of the cloud-belts across the disk; and more recent observation with the spectroscope seems to prove that Jupiter emits light from its surface analogous to that of the red stars. The conspicuous flattening of its poles is another feature that science would expect in a rapidly rotating liquid globe. In a word, Jupiter seems to be in the last stage of stellar development. Such, at some remote time, was our earth; such one day will be the sun.

The neighbouring planet Saturn supports the conclusion. Here again we have a gigantic globe, 28,000 miles in diameter, turning on its axis in the short space of ten hours; and here again we find the conspicuous flattening of the poles, the trailing belts of massed vapour across the disk, the red glow lighting the edges of the belts, and the spectroscopic evidence of an emission of light. Once more it is difficult to doubt that a highly heated body is wrapped in that thick mantle of vapour. With its ten moons and its marvellous ring-system--an enormous collection of fragments, which the influence of the planet or of its nearer satellites seems to have prevented from concentrating--Saturn has always been a beautiful object to observe; it is not less interesting in those features which we faintly detect in its disk.

The next planet, Uranus, 32,000 miles in diameter, seems to be another cloud-wrapt, greatly heated globe, if not, as some think, a sheer mass of vapours without a liquid core. Neptune is too dim and distant for profitable examination. It may be added, however, that the dense masses of gas which are found to surround the outer planets seem to confirm the nebular theory, which assumes that they were developed in the outer and lighter part of the material hurled from the sun.

From this encouraging survey of the sister-planets we return with more confidence to the story of the earth. I will not attempt to follow an imaginative scheme in regard to its early development. Take four photographs --one of a spiral nebula without knots in its arms, one of a nebula like that in Canes Venatici, one of the sun, and one of Jupiter--and you have an excellent illustration of the chief stages in its formation. In the first picture a section of the luminous arm of the nebula stretches thinly across millions of miles of space. In the next stage this material is largely collected in a luminous and hazy sphere, as we find in the nebula in Canes Venatici. The sun serves to illustrate a further stage in the condensation of this sphere. Jupiter represents a later chapter, in which the cooler vapours are wrapped close about the red-hot body of the planet. That seems to have been the early story of the earth. Some 6,000,000,000 billion tons of the nebulous matter were attracted to a common centre. As the particles pressed centreward, the temperature rose, and for a time the generation of heat was greater than its dissipation. Whether the earth ever shone as a small white star we cannot say. We must not hastily conclude that such a relatively small mass would behave like the far greater mass of a star, but we may, without attempting to determine its temperature, assume that it runs an analogous course.

One of the many features which I have indicated as pointing to a former fluidity of the earth may be explained here. We shall see in the course of this work that the mountain chains and other great irregularities of the earth's surface appear at a late stage in its development. Even as we find them to-day, they are seen to be merely slight ridges and furrows on the face of the globe, when we reflect on its enormous diameter, but there is good reason to think that in the beginning the earth was much nearer to a perfectly globular form. This points to a liquid or gaseous condition at one time, and the flattening of the sphere at the poles confirms the impression. We should hardly expect so perfect a rotundity in a body formed by the cool accretion of solid fragments and particles. It is just what we should expect in a fluid body, and the later irregularities of the surface are accounted for by the constant crumpling and wearing of its solid crust. Many would find a confirmation of this in the phenomena of volcanoes, geysers, and earthquakes, and the increase of the temperature as we descend the crust. But the interior condition of the earth, and the nature of these phenomena, are much disputed at present, and it is better not to rely on any theory of them. It is suggested that radium may be responsible for this subterraneous heat.

The next stage in the formation of the earth is necessarily one that we can reach only by conjecture. Over the globe of molten fire the vapours and gases would be suspended like a heavy canopy, as we find in Jupiter and Saturn to-day. When the period of maximum heat production was passed, however, the radiation into space would cause a lowering of the temperature, and a scum would form on the molten surface. As may be observed on the surface of any cooling vessel of fluid, the scum would stretch and crack; the skin would, so to say, prove too small for the body. The molten ocean below would surge through the crust, and bury it under floods of lava. Some hold that the slabs would sink in the ocean of metal, and thus the earth would first solidify in its deeper layers. There would, in any case, be an age-long struggle between the molten mass and the confining crust, until at length--to employ the old Roman conception of the activity of Etna--the giant was imprisoned below the heavy roof of rock.

Here again we seem to find evidence of the general correctness of the theory. The objection has been raised that the geologist does not find any rocks which he can identify as portions of the primitive crust of the earth. It seems to me that it would be too much to expect the survival at the surface of any part of the first scum that cooled on that fiery ocean. It is more natural to suppose that millions of years of volcanic activity on a prodigious scale would characterise this early stage, and the "primitive crust" would be buried in fragments, or dissolved again, under deep seas of lava. Now, this is precisely what we find, The oldest rocks known to the geologist--the Archaean rocks--are overwhelmingly volcanic, especially in their lower part. Their thickness, as we know them, is estimated at 50,000 feet; a thickness which must represent many millions of years. But we do not know how much thicker than this they may be. They underlie the oldest rocks that have ever been exposed to the gaze of the geologist. They include sedimentary deposits, showing the action of water, and even probable traces of organic remains, but they are, especially in their deeper and older sections, predominantly volcanic. They evince what we may call a volcanic age in the early story of the planet.

But before we pursue this part of the story further we must interpolate a remarkable event in the record--the birth of the moon. It is now generally believed, on a theory elaborated by Sir G. Darwin, that when the formation of the crust had reached a certain depth--something over thirty miles, it is calculated--it parted with a mass of matter, which became the moon. The size of our moon, in comparison with the earth, is so exceptional among the satellites which attend the planets of our solar system that it is assigned an exceptional origin. It is calculated that at that time the earth turned on its axis in the space of four or five hours, instead of twenty-four. We have already seen that the tidal influence of the sun has the effect of moderating the rotation of the planets. Now, this very rapid rotation of a liquid mass, with a thin crust, would (together with the instability occasioned by its cooling) cause it to bulge at the equator. The bulge would increase until the earth became a pear-shaped body. The small end of the pear would draw further and further away from the rest--as a drop of water does on the mouth of a tap--and at last the whole mass (some 5,000,000,000 cubic miles of matter) was broken off, and began to pursue an independent orbit round the earth.

There are astronomers who think that other cosmic bodies, besides our moon, may have been formed in this way. Possibly it is true of some of the double stars, but we will not return to that question. The further story of the moon, as it is known to astronomers, may be given in a few words. The rotational movement of the earth is becoming gradually slower on account of tidal influence; our day, in fact, becomes an hour longer every few million years. It can be shown that this had the effect of increasing the speed, and therefore enlarging the orbit, of the moon, as it revolved round the earth. As a result, the moon drew further and further away from the earth until it reached its present position, about 240,000 miles away. At the same time the tidal influence of the earth was lessening the rotational movement of the moon. This went on until it turned on its axis in the same period in which it revolves round the earth, and on this account it always presents the same face to the earth.

Through what chapters of life the moon may have passed in the meantime it is impossible to say. Its relatively small mass may have been unable to keep the lighter gases at its surface, or its air and water may, as some think, have been absorbed. It is to-day practically an airless and waterless desert, alternating between the heat of its long day and the intense cold of its long night. Careful observers, such as Professor Pickering, think that it may still have a shallow layer of heavy gases at its surface, and that this may permit the growth of some stunted vegetation during the day. Certain changes of colour, which are observed on its surface, have been interpreted in that sense. We can hardly conceive any other kind of life on it. In the dark even the gases will freeze on its surface, as there is no atmosphere to retain the heat. Indeed, some students of the moon (Fauth, etc.) believe that it is an unchanging desert of ice, bombarded by the projectiles of space.

An ingenious speculation as to the effect on the earth of this dislodgment of 5,000,000,000 cubic miles of its substance is worth noting. It supposes that the bed of the Pacific Ocean represents the enormous gap torn in its side by the delivery of the moon. At each side of this chasm the two continents, the Old World and the New, would be left floating on their molten ocean; and some have even seen a confirmation of this in the lines of crustal weakness which we trace, by volcanoes and earthquakes, on either side of the Pacific. Others, again, connect the shape of our great masses of land, which generally run to a southern point, with this early catastrophe. But these interesting speculations have a very slender basis, and we will return to the story of the development of the earth.

The last phase in preparation for the appearance of life would be the formation of the ocean. On the lines of the generally received nebular hypothesis this can easily be imagined, in broad outline. The gases would form the outer shell of the forming planet, since the heavier particles would travel inward. In this mixed mass of gas the oxygen and hydrogen would combine, at a fitting temperature, and form water. For ages the molten crust would hold this water suspended aloft as a surrounding shell of cloud, but when the surface cooled to about 380 degrees C. (Sollas), the liquid would begin to pour on it. A period of conflict would ensue, the still heated crust and the frequent volcanic outpours sending the water back in hissing steam to the clouds. At length, and now more rapidly, the temperature of the crust would sink still lower, and a heated ocean would settle upon it, filling the hollows of its irregular surface, and washing the bases of its outstanding ridges. From that time begins the age-long battle of the land and the water which, we shall see, has had a profound influence on the development of life.

In deference to the opinion of a number of geologists we must glance once more at the alternative view of the planetesimal school. In their opinion the molecules of water were partly attracted to the surface out of the disrupted matter, and partly collected within the porous outer layers of the globe. As the latter quantity grew, it would ooze upwards, fill the smaller depressions in the crust, and at length, with the addition of the attracted water, spread over the irregular surface. There is an even more important difference of opinion in regard to the formation of the atmosphere, but we may defer this until the question of climate interests us. We have now made our globe, and will pass on to that early chapter of its story in which living things make their appearance.

To some it will seem that we ought not to pass from the question of origin without a word on the subject of the age of the earth. All that one can do, however, is to give a number of very divergent estimates. Physicists have tried to calculate the age of the sun from the rate of its dissipation of heat, and have assigned, at the most, a hundred million years to our solar system; but the recent discovery of a source of heat in the disintegration of such metals as radium has made their calculations useless. Geologists have endeavoured, from observation of the action of geological agencies to-day, to estimate how long it will have taken them to form the stratified crust of the earth; but even the best estimates vary between twenty-five and a hundred million years, and we have reason to think that the intensity of these geological agencies may have varied in different ages. Chemists have calculated how long it would take the ocean, which was originally fresh water, to take up from the rocks and rivers the salt which it contains to-day; Professor Joly has on this ground assigned a hundred million years since the waters first descended upon the crust. We must be content to know that the best recent estimates, based on positive data, vary between fifty and a hundred million years for the story which we are now about to narrate. The earlier or astronomical period remains quite incalculable. Sir G. Darwin thinks that it was probably at least a thousand million years since the moon was separated from the earth. Whatever the period of time may be since some cosmic cataclysm scattered the material of our solar system in the form of a nebula, it is only a fraction of that larger and illimitable time which the evolution of the stars dimly suggests to the scientific imagination.

THE GEOLOGICAL SERIES

[The scale of years adopted--50,000,000 for the stratified rocks--is merely an intermediate between conflicting estimates.]

ERA. PERIOD. RELATIVE LENGTH.

Quaternary Holocene 500,000 years Pleistocene
Tertiary Pliocene 5,500,000 years or Miocene Cenozoic Oligocene Eocene
Secondary Cretaceous 7,200,000 years or Jurassic 3,600,000 " Mesozoic Triassic 2,500,000 "
Primary Permian 2,800,000 years or Carboniferous 6,200,000 " Palaeozoic Devonian 8,000,000 " Silurian 5,400,000 " Ordovician 5,400,000 " Cambrian 8,000,000 "
Archaean Keweenawan Unknown (probably Animikie at least Huronian 50,000,000 years) Keewatin Laurentian

CHAPTER V. THE BEGINNING OF LIFE

There is, perhaps, no other chapter in the chronicle of the earth that we approach with so lively an interest as the chapter which should record the first appearance of life. Unfortunately, as far as the authentic memorials of the past go, no other chapter is so impenetrably obscure as this. The reason is simple. It is a familiar saying that life has written its own record, the long-drawn record of its dynasties and its deaths, in the rocks. But there were millions of years during which life had not yet learned to write its record, and further millions of years the record of which has been irremediably destroyed. The first volume of the geological chronicle of the earth is the mass of the Archaean (or "primitive") rocks. What the actual magnitude of that volume, and the span of time it covers, may be, no geologist can say. The Archaean rocks still solidly underlie the lowest depth he has ever reached. It is computed, however, that these rocks, as far as they are known to us, have a total depth of nearly ten miles, and seem therefore to represent at least half the story of the earth from the time when it rounded into a globe, or cooled sufficiently to endure the presence of oceans.

Yet all that we read of the earth's story during those many millions of years could be told in a page or two. That section of geology is still in its infancy, it is true. A day may come when science will decipher a long and instructive narrative in the masses of quartz and gneiss, and the layers of various kinds, which it calls the Archaean rocks. But we may say with confidence that it will not discover in them more than a few stray syllables of the earlier part, and none whatever of the earliest part, of the epic of living nature. A few fossilised remains of somewhat advanced organisms, such as shell-fish and worms, are found in the higher and later rocks of the series, and more of the same comparatively high types will probably appear. In the earlier strata, representing an earlier stage of life, we find only thick seams of black shale, limestone, and ironstone, in which we seem to see the ashes of primitive organisms, cremated in the appalling fires of the volcanic age, or crushed out of recognition by the superimposed masses. Even if some wizardry of science were ever to restore the forms that have been reduced to ashes in this Archaean crematorium, it would be found that they are more or less advanced forms, far above the original level of life. No trace will ever be found in the rocks of the first few million years in the calendar of life.

The word impossible or unknowable is not lightly uttered in science to-day, but there is a very plain reason for admitting it here. The earliest living things were at least as primitive of nature as the lowest animals and plants we know to-day, and these, up to a fair level of organisation, are so soft of texture that, when they die, they leave no remains which may one day be turned into fossils. Some of them, indeed, form tiny shells of flint or lime, or, like the corals, make for themselves a solid bed; but this is a relatively late and higher stage of development. Many thousands of species of animals and plants lie below that level. We are therefore forced to conclude, from the aspect of living nature to-day, that for ages the early organisms had no hard and preservable parts. In thus declaring the impotence of geology, however, we are at the same time introducing another science, biology, which can throw appreciable light on the evolution of life. Let us first see what geology tells us about the infancy of the earth.

The distribution of the early rocks suggests that there was comparatively little dry land showing above the surface of the Archaean ocean. Our knowledge of these rocks is not at all complete, and we must remember that some of this primitive land may be now under the sea or buried in unsuspected regions. It is significant, however, that, up to the present, exploration seems to show that in those remote ages only about one-fifth of our actual land-surface stood above the level of the waters. Apart from a patch of some 20,000 square miles of what is now Australia, and smaller patches in Tasmania, New Zealand, and India, nearly the whole of this land was in the far North. A considerable area of eastern Canada had emerged, with lesser islands standing out to the west and south of North America. Another large area lay round the basin of the Baltic; and as Greenland, the Hebrides, and the extreme tip of Scotland, belong to the same age, it is believed that a continent, of which they are fragments, united America and Europe across the North Atlantic. Of the rest of what is now Europe there were merely large islands--one on the border of England and Wales, others in France, Spain, and Southern Germany. Asia was represented by a large area in China and Siberia, and an island or islands on the site of India. Very little of Africa or South America existed.

It will be seen at a glance that the physical story of the earth from that time is a record of the emergence from the waters of larger continents and the formation of lofty chains of mountains. Now this world-old battle of land and sea has been waged with varying fortune from age to age, and it has been one of the most important factors in the development of life. We are just beginning to realise what a wonderful light it throws on the upward advance of animals and plants. No one in the scientific world to-day questions that, however imperfect the record may be, there has been a continuous development of life from the lowest level to the highest. But why there was advance at all, why the primitive microbe climbs the scale of being, during millions of years, until it reaches the stature of humanity, seems to many a profound mystery. The solution of this mystery begins to break upon us when we contemplate, in the geological record, the prolonged series of changes in the face of the earth itself, and try to realise how these changes must have impelled living things to fresh and higher adaptations to their changing surroundings.

Imagine some early continent with its population of animals and plants. Each bay, estuary, river, and lake, each forest and marsh and solid plain, has its distinctive inhabitants. Imagine this continent slowly sinking into the sea, until the advancing arms of the salt water meet across it, mingling their diverse populations in a common world, making the fresh-water lake brackish or salt, turning the dry land into swamp, and flooding the forest. Or suppose, on the other hand, that the land rises, the marsh is drained, the genial climate succeeded by an icy cold, the luscious vegetation destroyed, the whole animal population compelled to change its habits and its food. But this is no imaginary picture. It is the actual story of the earth during millions of years, and it is chiefly in the light of these vast and exacting changes in the environment that we are going to survey the panorama of the advance of terrestrial life.

For the moment it will be enough to state two leading principles. The first is that there is no such thing as a "law of evolution" in the sense in which many people understand that phrase. It is now sufficiently well known that, when science speaks of a law, it does not mean that there is some rule that things MUST act in such and such a way. The law is a mere general expression of the fact that they DO act in that way. But many imagine that there is some principle within the living organism which impels it onward to a higher level of organisation. That is entirely an error. There is no "law of progress." If an animal is fitted to secure its livelihood and breed posterity in certain surroundings, it may remain unchanged indefinitely if these surroundings do not materially change. So the duckmole of Australia and the tuatara of New Zealand have retained primitive features for millions of years; so the aboriginal Australian and the Fuegian have remained stagnant, in their isolation, for a hundred thousand years or more; so the Chinaman, in his geographical isolation, has remained unchanged for two thousand years. There is no more a "conservative instinct" in Chinese than there is a "progressive instinct" in Europeans. The difference is one of history and geography, as we shall see.

To make this important principle still clearer, let us imagine some primitive philosopher observing the advance of the tide over a level beach. He must discover two things: why the water comes onward at all, and why it advances along those particular channels. We shall see later how men of science explain or interpret the mechanism in a living thing which enables it to advance, when it does advance. For the present it is enough to say that new-born animals and plants are always tending to differ somewhat from their parents, and we now know, by experiment, that when some exceptional influence is brought to bear on the parent, the young may differ considerably from her. But, if the parents were already in harmony with their environment, these variations on the part of the young are of no consequence. Let the environment alter, however, and some of these variations may chance to make the young better fitted than the parent was. The young which happen to have the useful variation will have an advantage over their brothers or sisters, and be more likely to survive and breed the next generation. If the change in the environment (in the food or climate, for instance) is prolonged and increased for hundreds of thousands of years, we shall expect to find a corresponding change in the animals and plants.

We shall find such changes occurring throughout the story of the earth. At one important point in the story we shall find so grave a revolution in the face of nature that twenty-nine out of every thirty species of animals and plants on the earth are annihilated. Less destructive and extreme changes have been taking place during nearly the whole of the period we have to cover, entailing a more gradual alteration of the structure of animals and plants; but we shall repeatedly find them culminating in very great changes of climate, or of the distribution of land and water, which have subjected the living population of the earth to the most searching tests and promoted every variation toward a more effective organisation.*

* This is a very simple expression of "Darwinism," and will be enlarged later. The reader should ignore the occasional statement of non-scientific writers that Darwinism is "dead" or superseded. The questions which are actually in dispute relate to the causes of the variation of the young from their parents, the magnitude of these variations' and the transmission of changes acquired by an animal during its own life. We shall see this more fully at a later stage. The importance of the environment as I have described it, is admitted by all schools.



And the second guiding principle I wish to lay down in advance is that these great changes in the face of the earth, which explain the progress of organisms, may very largely be reduced to one simple agency--the battle of the land and the sea. When you gaze at some line of cliffs that is being eaten away by the waves, or reflect on the material carried out to sea by the flooded river, you are--paradoxical as it may seem--beholding a material process that has had a profound influence on the development of life. The Archaean continent that we described was being reduced constantly by the wash of rain, the scouring of rivers, and the fretting of the waves on the coast. It is generally thought that these wearing agencies were more violent in early times, but that is disputed, and we will not build on it. In any case, in the course of time millions of tons of matter were scraped off the Archaean continent and laid on the floor of the sea by its rivers. This meant a very serious alteration of pressure or weight on the surface of the globe, and was bound to entail a reaction or restoration of the balance.

The rise of the land and formation of mountains used to be ascribed mainly to the cooling and shrinking of the globe of the earth. The skin (crust), it was thought, would become too large for the globe as it shrank, and would wrinkle outwards, or pucker up into mountain-chains. The position of our greater mountain-chains sprawling across half the earth (the Pyrenees to the Himalaya, and the Rocky Mountains to the Andes), seems to confirm this, but the question of the interior of the earth is obscure and disputed, and geologists generally conceive the rise of land and formation of mountains in a different way. They are due probably to the alteration of pressure on the crust in combination with the instability of the interior. The floors of the seas would sink still lower under their colossal burdens, and this would cause some draining of the land-surface. At the same time the heavy pressure below the seas and the lessening of pressure over the land would provoke a reaction. Enormous masses of rock would be forced toward and underneath the land-surface, bending, crumpling, and upheaving it as if its crust were but a leather coat. As a result, masses of land would slowly rise above the plain, to be shaped into hills and valleys by the hand of later time, and fresh surfaces would be dragged out of the deep, enlarging the fringes of the primitive continents, to be warped and crumpled in their turn at the next era of pressure.

In point of geological fact, the story of the earth has been one prolonged series of changes in the level of land and water, and in their respective limits. These changes have usually been very gradual, but they have always entailed changes (in climate, etc. ) of the greatest significance in the evolution of life. What was the swampy soil of England in the Carboniferous period is now sometimes thousands of feet beneath us; and what was the floor of a deep ocean over much of Europe and Asia at another time is now to be found on the slopes of lofty Alps, or 20,000 feet above the sea-level in Thibet. Our story of terrestrial life will be, to a great extent, the story of how animals and plants changed their structure in the long series of changes which this endless battle of land and sea brought over the face of the earth.

As we have no recognisable remains of the animals and plants of the earliest age, we will not linger over the Archaean rocks. Starting from deep and obscure masses of volcanic matter, the geologist, as he travels up the series of Archaean rocks, can trace only a dim and most unsatisfactory picture of those remote times. Between outpours of volcanic floods he finds, after a time, traces that an ocean and rivers are wearing away the land. He finds seams of carbon among the rocks of the second division of the Archaean (the Keewatin), and deduces from this that a dense sea-weed population already covered the floor of the ocean. In the next division (the Huronian) he finds the traces of extensive ice-action strangely lying between masses of volcanic rock, and sees that thousands of square miles of eastern North America were then covered with an ice-sheet. Then fresh floods of molten matter are poured out from the depths below; then the sea floods the land for a time; and at last it makes its final emergence as the first definitive part of the North American continent, to enlarge, by successive fringes, to the continent of to-day.*

* I am quoting Professor Coleman's summary of Archaean research in North America (Address to the Geological Section of the British Association, 1909). Europe, as a continent, has had more "ups and downs" than America in the course of geological time.


This meagre picture of the battle of land and sea, with interludes of great volcanic activity and even of an ice age, represents nearly all we know of the first half of the world's story from geology. It is especially disappointing in regard to the living population. The very few fossils we find in the upper Archaean rocks are so similar to those we shall discuss in the next chapter that we may disregard them, and the seams of carbon-shales, iron-ore, and limestone, suggest only, at the most, that life was already abundant. We must turn elsewhere for some information on the origin and early development of life.

The question of the origin of life I will dismiss with a brief account of the various speculations of recent students of science. Broadly speaking, their views fall into three classes. Some think that the germs of life may have come to the earth from some other body in the universe; some think that life was evolved out of non-living matter in the early ages of the earth, under exceptional conditions which we do not at present know, or can only dimly conjecture; and some think that life is being evolved from non-life in nature to-day, and always has been so evolving. The majority of scientific men merely assume that the earliest living things were no exception to the general process of evolution, but think that we have too little positive knowledge to speculate profitably on the manner of their origin.

The first view, that the germs of life may have come to this planet on a meteoric visitor from some other world, as a storm-driven bird may take its parasites to some distant island, is not without adherents to-day. It was put forward long ago by Lord Kelvin and others; it has been revived by the distinguished Swede, Professor Svante Arrhenius. The scientific objection to it is that the more intense (ultra-violet) rays of the sun would frill such germs as they pass through space. But a broader objection, and one that may dispense us from dwelling on it, is that we gain nothing by throwing our problems upon another planet. We have no ground for supposing that the earth is less capable of evolving life than other planets.

The second view is that, when the earth had passed through its white-hot stage, great masses of very complex chemicals, produced by the great heat, were found on its surface. There is one complex chemical substance in particular, called cyanogen, which is either an important constituent of living matter, or closely akin to it. Now we need intense heat to produce this substance in the laboratory. May we not suppose that masses of it were produced during the incandescence of the earth, and that, when the waters descended, they passed through a series of changes which culminated in living plasm? Such is the "cyanogen hypothesis" of the origin of life, advocated by able physiologists such as Pfluger, Verworn, and others. It has the merit of suggesting a reason why life may not be evolving from non-life in nature to-day, although it may have so evolved in the Archaean period.

Other students suggest other combinations of carbon-compounds and water in the early days. Some suggest that electric action was probably far more intense in those ages; others think that quantities of radium may have been left at the surface. But the most important of these speculations on the origin of life in early times, and one that has the merit of not assuming any essentially different conditions then than we find now, is contained in a recent pronouncement of one of the greatest organic chemists in Europe, Professor Armstrong. He says that such great progress has been made in his science--the science of the chemical processes in living things--that "their cryptic character seems to have disappeared almost suddenly." On the strength of this new knowledge of living matter, he ventures to say that "a series of lucky accidents" could account for the first formation of living things out of non-living matter in Archaean times. Indeed, he goes further. He names certain inorganic substances, and says that the blowing of these into pools by the wind on the primitive planet would set afoot chemical combinations which would issue in the production of living matter.*

* See his address in Nature, vol. 76, p. 651. For other speculations see Verworn's "General Physiology," Butler Burke's "Origin of Life" (1906), and Dr. Bastian's "Origin of Life" (1911).


It is evident that the popular notion that scientific men have declared that life cannot be evolved from non-life is very far astray. This blunder is usually due to a misunderstanding of the dogmatic statement which one often reads in scientific works that "every living thing comes from a living thing." This principle has no reference to remote ages, when the conditions may have been different. It means that to-day, within our experience, the living thing is always born of a living parent. However, even this is questioned by some scientific men of eminence, and we come to the third view.

Professor Nageli, a distinguished botanist, and Professor Haeckel, maintain that our experience, as well as the range of our microscopes, is too limited to justify the current axiom. They believe that life may be evolving constantly from inorganic matter. Professor J. A. Thomson also warns us that our experience is very limited, and, for all we know, protoplasm may be forming naturally in our own time. Mr. Butler Burke has, under the action of radium, caused the birth of certain minute specks which strangely imitate the behaviour of bacteria. Dr. Bastian has maintained for years that he has produced living things from non-living matter. In his latest experiments, described in the book quoted, purely inorganic matter is used, and it is previously subjected, in hermetically sealed tubes, to a heat greater than what has been found necessary to kill any germs whatever.

Evidently the problem of the origin of life is not hopeless, but our knowledge of the nature of living matter is still so imperfect that we may leave detailed speculation on its origin to a future generation. Organic chemistry is making such strides that the day may not be far distant when living matter will be made by the chemist, and the secret of its origin revealed. For the present we must be content to choose the more plausible of the best-informed speculations on the subject.

But while the origin of life is obscure, the early stages of its evolution come fairly within the range of our knowledge. To the inexpert it must seem strange that, whereas we must rely on pure speculation in attempting to trace the origin of life, we can speak with more confidence of those early developments of plants and animals which are equally buried in the mists of the Archaean period. Have we not said that nothing remains of the procession of organisms during half the earth's story but a shapeless seam of carbon or limestone?

A simple illustration will serve to justify the procedure we are about to adopt. Suppose that the whole of our literary and pictorial references to earlier stages in the development of the bicycle, the locomotive, or the loom, were destroyed. We should still be able to retrace the phases of their evolution, because we should discover specimens belonging to those early phases lingering in our museums, in backward regions, and elsewhere. They might yet be useful in certain environments into which the higher machines have not penetrated. In the same way, if all the remains of prehistoric man and early civilisation were lost, we could still fairly retrace the steps of the human race, by gathering the lower tribes and races, and arranging them in the order of their advancement. They are so many surviving illustrations of the stages through which mankind as a whole has passed.

Just in the same way we may marshal the countless species of animals and plants to-day in such order that they will, in a general way, exhibit to us the age-long procession of life. From the very start of living evolution certain forms dropped out of the onward march, and have remained, to our great instruction, what their ancestors were millions of years ago. People create a difficulty for themselves by imagining that, if evolution is true, all animals must evolve. A glance at our own fellows will show the error of this. Of one family of human beings, as a French writer has said, one only becomes a Napoleon; the others remain Lucien, Jerome, or Joseph. Of one family of animals or trees, some advance in one or other direction; some remain at the original level. There is no "law of progress." The accidents of the world and hereditary endowment impel some onward, and do not impel others. Hence at nearly every great stage in the upward procession through the ages some regiment of plants or animals has dropped out, and it represents to-day the stage of life at which it ceased to progress. In other words, when we survey the line of the hundreds of thousands of species which we find in nature to-day, we can trace, amid their countless variations and branches, the line of organic evolution in the past; just as we could, from actual instances, study the evolution of a British house, from the prehistoric remains in Devonshire to a mansion in Park Lane or a provincial castle.

Another method of retracing the lost early chapters in the development of life is furnished by embryology. The value of this method is not recognised by all embryologists, but there are now few authorities who question the substantial correctness of it, and we shall, as we proceed, see some remarkable applications of it. In brief, it is generally admitted that an animal or plant is apt to reproduce, during its embryonic development, some of the stages of its ancestry in past time. This does not mean that a higher animal, whose ancestors were at one time worms, at another time fishes, and at a later time reptiles, will successively take the form of a little worm, a little fish, and a little reptile. The embryonic life itself has been subject to evolution, and this reproduction of ancestral forms has been proportionately disturbed. Still, we shall find that animals will tend, in their embryonic development, to reproduce various structural features which can only be understood as reminiscences of ancestral organs. In the lower animals the reproduction is much less disturbed than in the higher, but even in the case of man this law is most strikingly verified. We shall find it useful sometimes at least in confirming our conclusions as to the ancestry of a particular group.

We have, therefore, two important clues to the missing chapters in the story of evolution. Just as the scheme of the evolution of worlds is written broadly across the face of the heavens to-day, so the scheme of the evolution of life is written on the face of living nature; and it is written again, in blurred and broken characters, in the embryonic development of each individual. With these aids we set out to restore the lost beginning of the epic of organic evolution.





CHAPTER VI. THE INFANCY OF THE EARTH

The long Archaean period, into which half the story of the earth is so unsatisfactorily packed, came to a close with a considerable uplift of the land. We have seen that the earth at times reaches critical stages owing to the transfer of millions of tons of matter from the land to the depths of the ocean, and the need to readjust the pressure on the crust. Apparently this stage is reached at the end of the Archaean, and a great rise of the land --probably protracted during hundreds of thousands of years--takes place. The shore-bottoms round the primitive continent are raised above the water, their rocks crumpling like plates of lead under the overpowering pressure. The sea retires with its inhabitants, mingling their various provinces, transforming their settled homes. A larger continent spans the northern ocean of the earth.

In the shore-waters of this early continent are myriads of living things, representing all the great families of the animal world below the level of the fish and the insect. The mud and sand in which their frames are entombed, as they die, will one day be the "Cambrian" rocks of the geologist, and reveal to him their forms and suggest their habits. No great volcanic age will reduce them to streaks of shapeless carbon. The earth now buries its dead, and from their petrified remains we conjure up a picture of the swarming life of the Cambrian ocean.

A strange, sluggish population burrows in the mud, crawls over the sand, adheres to the rocks, and swims among the thickets of sea-weed. The strangest and most formidable, though still too puny a thing to survive in a more strenuous age, is the familiar Trilobite of the geological museum; a flattish animal with broad, round head, like a shovel, its back covered with a three-lobed shell, and a number of fine legs or swimmers below. It burrows in the loose bottom, or lies in it with its large compound eyes peeping out in search of prey. It is the chief representative of the hard-cased group (Crustacea) which will later replace it with the lobster, the shrimp, the crab, and the water-flea. Its remains form from a third to a fourth of all the buried Cambrian skeletons. With it, swimming in the water, are smaller members of the same family, which come nearer to our familiar small Crustacea.

Shell-fish are the next most conspicuous inhabitants. Molluscs are already well represented, but the more numerous are the more elementary Brachiopods ("lampshells"), which come next to the Trilobites in number and variety. Worms (or Annelids) wind in and out of the mud, leaving their tracks and tubes for later ages. Strange ball or cup-shaped little animals, with a hard frame, mounted on stony stalks and waving irregular arms to draw in the food-bearing water, are the earliest representatives of the Echinoderms. Some of these Cystids will presently blossom into the wonderful sea-lily population of the next age, some are already quitting their stalks, to become the free-moving star-fish, of which a primitive specimen has been found in the later Cambrian. Large jelly-fishes (of which casts are preserved) swim in the water; coral-animals lay their rocky foundations, but do not as yet form reefs; coarse sponges rise from the floor; and myriads of tiny Radiolaria and Thalamophores, with shells of flint and lime, float at the surface or at various depths.

This slight sketch of the Cambrian population shows us that living things had already reached a high level of development. Their story evidently goes back, for millions of years, deep into those mists of the Archaean age which we were unable to penetrate. We turn therefore to the zoologist to learn what he can tell us of the origin and family-relations of these Cambrian animals, and will afterwards see how they are climbing to higher levels under the eye of the geologist.

At the basis of the living world of to-day is a vast population of minute, generally microscopic, animals and plants, which are popularly known as "microbes." Each consists, in scientific language, of one cell. It is now well known that the bodies of the larger animals and plants are made up of millions of these units of living matter, or cells--the atoms of the organic world--and I need not enlarge on it. But even a single cell lends itself to infinite variety of shape, and we have to penetrate to the very lowest level of this luxuriant world of one-celled organisms to obtain some idea of the most primitive living things. Properly speaking, there were no "first living things." It cannot be doubted by any student of nature that the microbe developed so gradually that it is as impossible to fix a precise term for the beginning of life as it is to say when the night ends and the day begins. In the course of time little one-celled living units appeared in the waters of the earth, whether in the shallow shore waters or on the surface of the deep is a matter of conjecture.

We are justified in concluding that they were at least as rudimentary in structure and life as the lowest inhabitants of nature to-day. The distinction of being the lowest known living organisms should, I think, be awarded to certain one-celled vegetal organisms which are very common in nature. Minute simple specks of living matter, sometimes less than the five-thousandth of an inch in diameter, these lowly Algae are so numerous that it is they, in their millions, which cover moist surfaces with the familiar greenish or bluish coat. They have no visible organisation, though, naturally, they must have some kind of structure below the range of the microscope. Their life consists in the absorption of food-particles, at any point of their surface, and in dividing into two living microbes, instead of dying, when their bulk increases. A very lowly branch of the Bacteria (Nitrobacteria) sometimes dispute their claim to the lowest position in the hierarchy of living nature, but there is reason to suspect that these Bacteria may have degenerated from a higher level.

Here we have a convenient starting-point for the story of life, and may now trace the general lines of upward development. The first great principle to be recognised is the early division of these primitive organisms into two great classes, the moving and the stationary. The clue to this important divergence is found in diet. With exceptions on both sides, we find that the non-moving microbes generally feed on inorganic matter, which they convert into plasm; the moving microbes generally feed on ready-made plasm--on the living non-movers, on each other, or on particles of dead organic matter. Now, inorganic food is generally diffused in the waters, so that the vegetal feeders have no incentive to develop mobility. On the other hand, the power to move in search of their food, which is not equally diffused, becomes a most important advantage to the feeders on other organisms. They therefore develop various means of locomotion. Some flow or roll slowly along like tiny drops of oil on an inclined surface; others develop minute outgrowths of their substance, like fine hairs, which beat the water as oars do. Some of them have one strong oar, like the gondolier (but in front of the boat); others have two or more oars; while some have their little flanks bristling with fine lashes, like the flanks of a Roman galley.

If we imagine this simple principle at work for ages among the primitive microbes, we understand the first great division of the living world, into plants and animals. There must have been a long series of earlier stages below the plant and animal. In fact, some writers insist that the first organisms were animal in nature, feeding on the more elementary stages of living matter. At last one type develops chlorophyll (the green matter in leaves), and is able to build up plasm out of inorganic matter; another type develops mobility, and becomes a parasite on the plant world. There is no rigid distinction of the two worlds. Many microscopic plants move about just as animals do, and many animals live on fixed stalks; while many plants feed on organic matter. There is so little "difference of nature" between the plant and the animal that the experts differ in classifying some of these minute creatures. In fact, we shall often find plants and animals crossing the line of division. We shall find animals rooting themselves to the floor, like plants, though they will generally develop arms or streamers for bringing the food to them; and we shall find plants becoming insect-catchers. All this merely shows that the difference is a natural tendency, which special circumstances may overrule. It remains true that the great division of the organic world is due to a simple principle of development; difference of diet leads to difference of mobility.

But this simple principle will have further consequences of a most important character. It will lead to the development of mind in one half of living nature and leave it undeveloped in the other. Mind, as we know it in the lower levels of life, is not confined to the animal at all. Many even of the higher plants are very delicately sensitive to stimulation, and at the lowest level many plants behave just like animals. In other words, this sensitiveness to stimuli, which is the first form of mind, is distributed according to mobility. To the motionless organism it is no advantage; to the pursuing and pursued organism it is an immense advantage, and is one of the chief qualities for natural selection to foster.

For the moment, however, we must glance at the operation of this and other natural principles in the evolution of the one-celled animals and plants, which we take to represent the primitive population of the earth. As there are tens of thousands of different species even of "microbes," it is clear that we must deal with them in a very summary way. The evolution of the plant I reserve for a later chapter, and I must be content to suggest the development of one-celled animals on very broad lines. When some of the primitive cells began to feed on each other, and develop mobility, it is probable that at least two distinct types were evolved, corresponding to the two lowest animal organisms in nature to-day. One of these is a very minute and very common (in vases of decaying flowers, for instance) speck of plasm, which moves about by lashing the water with a single oar (flagellum),

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